Can maternally transmitted endosymbionts facilitate the evolution of haplodiploidy?

J Evol Biol. 2006 Jan;19(1):194-202. doi: 10.1111/j.1420-9101.2005.00974.x.

Abstract

Whilst many invertebrate taxa are haplodiploid, the factors underlying the evolution of haplodiploidy remain unresolved. We investigate theoretically whether haplodiploidy might evolve as an outcome of the co-evolution between maternally inherited endosymbionts and their hosts. First, we substantially extend a recently developed model that involves maternally inherited endosymbionts that kill male offspring by eliminating the paternal genome. We also put forward a new hypothesis and develop a model that involves bacteria that induce cytoplasmic incompatibility (CI). Based on these models, we explore the co-evolutionary events that might occur between hosts and symbionts. We find that both with male-killers and CI-inducing endosymbionts, the hosts are likely to develop increased viability of haploid males, which can be considered a preadaptation to haplodiploidy. In addition, populations with haploidizing male-killers can in some cases evolve directly towards a genetic system of paternal genome elimination, a special form of haplodiploidy. These results are combined with consideration of mechanism and ecology to appraise the likelihood of male-killers and CI inducing bacteria being involved in the evolution of haplodiploidy.

Publication types

  • Comparative Study
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Biological Evolution*
  • Female
  • Invertebrates / genetics*
  • Invertebrates / microbiology*
  • Male
  • Models, Biological*
  • Ploidies*
  • Sex Ratio
  • Symbiosis*
  • Wolbachia / physiology