Pigeons were trained in a discrete-trial task in which a response to the center key obtained illumination of a side key and a single response to the side key terminated a trial with either reinforcement or nonreinforcement. Center-key speeds (i.e., reciprocals of latencies) declined with increases in intertrial interval, and it is argued that this effect is related to a decreased likelihood as intertrial interval increases that birds will be near the key at trial onset. Side-key speeds on trials with reinforcement decreased both with increases in intertrial interval and with shifts from continuous reinforcement to either a discrimination or a partial-reinforcement condition. The effects on side-key speeds are compared with effects observed in alley-running tasks using rats, and an interpretation in terms of frustration theory is offered for the results obtained in both types of task.