Abstract
The family 20 carbohydrate-binding module (CBM20) of the Arabidopsis starch phosphorylator glucan, water dikinase 3 (GWD3) was heterologously produced and its properties were compared to the CBM20 from a fungal glucoamylase (GA). The GWD3 CBM20 has 50-fold lower affinity for cyclodextrins than that from GA. Homology modelling identified possible structural elements responsible for this weak binding of the intracellular CBM20. Differential binding of fluorescein-labelled GWD3 and GA modules to starch granules in vitro was demonstrated by confocal laser scanning microscopy and yellow fluorescent protein-tagged GWD3 CBM20 expressed in tobacco confirmed binding to starch granules in planta.
Publication types
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Comparative Study
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Research Support, Non-U.S. Gov't
MeSH terms
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Arabidopsis / cytology
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Arabidopsis / enzymology*
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Arabidopsis / genetics
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Arabidopsis Proteins / genetics
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Arabidopsis Proteins / metabolism*
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Aspergillus niger / enzymology
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Aspergillus niger / genetics
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Cyclodextrins / chemistry
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Cytoplasmic Granules / genetics
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Cytoplasmic Granules / metabolism
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Fungal Proteins / genetics
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Fungal Proteins / metabolism
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Gene Expression
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Glucan 1,4-alpha-Glucosidase / genetics
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Glucan 1,4-alpha-Glucosidase / metabolism
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Microscopy, Confocal / methods
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Nicotiana / genetics
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Phosphotransferases (Paired Acceptors) / genetics
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Phosphotransferases (Paired Acceptors) / metabolism*
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Protein Binding / physiology
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Protein Structure, Tertiary / physiology
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Starch / genetics
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Starch / metabolism*
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Structural Homology, Protein
Substances
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Arabidopsis Proteins
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Cyclodextrins
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Fungal Proteins
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Starch
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Phosphotransferases (Paired Acceptors)
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alpha-glucan, water dikinase, Arabidopsis
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Glucan 1,4-alpha-Glucosidase