1 Department of Biology, MIT, Cambridge, MA 02139, Howard Hughes Medical Institute, MIT, Cambridge, MA 02139; Whitehead Institute for Biomedical Research, Cambridge MA 02142, Broad Institute of Harvard and MIT, Seven Cambridge Center, Cambridge, MA 02142, The David H. Koch Institute for Integrative Cancer Research at MIT, Cambridge, MA 02139, USA.
2 Department of Physiology, Institute for Diabetes, Obesity, and Metabolism, Perelman School of Medicine, University of Pennsylvania, Philadelphia PA 19104, USA.