1 Memorial Sloan Kettering Cancer Center, 1275 York Avenue, New York, NY 10065, USA.
2 Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, CT 06511, USA.
3 MRC London Institute of Medical Sciences, Hammersmith Campus, Du Cane Road, London W12 0HS, UK.
4 Faculty of Medicine, Imperial College London, Hammersmith Campus, Du Cane Road, London W12 0HS, UK.
5 Department of Biochemistry, University of Utah School of Medicine and Howard Hughes Medical Institute, 15 N. Medical Drive East, Salt Lake City, UT 84108, USA.