1 Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China.
2 National Key Laboratory of Wheat Improvement, College of Life Sciences, Shandong Agricultural University, Tai'an, China.
3 University of Chinese Academy of Sciences, Beijing, China.
5 National Key Laboratory of Wheat Improvement, College of Life Sciences, Shandong Agricultural University, Tai'an, China. zhangxs@sdau.edu.cn.
6 Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China. jxiao@genetics.ac.cn.
7 University of Chinese Academy of Sciences, Beijing, China. jxiao@genetics.ac.cn.
8 CAS-JIC Centre of Excellence for Plant and Microbial Science (CEPAMS), Institute of Genetics and Developmental Biology, CAS, Beijing, China. jxiao@genetics.ac.cn.