Quercetin attenuated necroptosis and apoptosis caused by LPS-induced mitochondrial function dysfunction through the METTL3-mediated PTEN m6A methylation/PI3K/AKT signaling in broiler livers

Phytomedicine. 2025 Apr:139:156551. doi: 10.1016/j.phymed.2025.156551. Epub 2025 Feb 21.

Abstract

Background: Quercetin (QUE), a natural flavonoid, offered an efficient protection against organism injury. N6-methyladenosine (m6A) methylation is considered to be the most prevalent and abundant modifications involved in various diseases.

Purpose: We sought to explore protective roles of QUE in mitigating necroptosis and apoptosis triggered by LPS-induced imbalances in mitochondria dynamic and energy metabolism in broiler livers, with a focus on m6A methylation modulation.

Study design/methods: We used LPS as a stimulus and treated with QUE to establish this in vivo and in vitro. In addition, we treated LMH cells with siMETTL3 (80 nM) to determine its detailed mechanism.

Results: Our findings revealed QUE significantly decreased METTL3 expression, leading to a decrease in PTEN m6A methylation and factors related to mitochondria fission, necroptosis, and apoptosis in the QUE+LPS group. In contrast, QUE treatment promoted the expression levels of marker factors for mitochondria fusion, energy metabolism, anti-apoptosis, and PI3K/AKT compared with the LPS group. Additionally, an increase of ΔΨm, ATP content, and ATPase activity was observed. AO/EB staining, Flow cytometry and TUNEL assays confirmed QUE inhibited LPS-induced apoptosis and necroptosis. Molecular docking analysis and cellular thermal shift assay supported an interaction between QUE and METTL3.

Conclusion: In summary, QUE mitigated necroptosis and apoptosis triggered by LPS-induced disorders of mitochondrial kinetic and metabolic processes in broiler livers through its interaction with METTL3, regulating PTEN m6A methylation/PI3K/AKT signaling pathway. This study enhances our understanding of biological functions for QUE and lays a theoretical foundation for developing new therapeutic interventions, highlighting its potential value.

Keywords: Broiler liver; Energy metabolism; Mitochondria dynamics; Necroptosis and apoptosis; PTEN m6A methylation; Quercetin.

MeSH terms

  • Adenosine / analogs & derivatives
  • Adenosine / metabolism
  • Animals
  • Apoptosis* / drug effects
  • Chickens
  • Lipopolysaccharides
  • Liver* / drug effects
  • Liver* / metabolism
  • Methylation / drug effects
  • Methyltransferases* / metabolism
  • Mitochondria* / drug effects
  • Mitochondria* / metabolism
  • Necroptosis* / drug effects
  • PTEN Phosphohydrolase / metabolism
  • Phosphatidylinositol 3-Kinases / metabolism
  • Proto-Oncogene Proteins c-akt / metabolism
  • Quercetin* / pharmacology
  • Signal Transduction / drug effects

Substances

  • Lipopolysaccharides
  • Methyltransferases
  • Quercetin
  • Proto-Oncogene Proteins c-akt
  • PTEN Phosphohydrolase
  • Phosphatidylinositol 3-Kinases
  • Adenosine
  • N-methyladenosine